When a new threat capable of destroying the entire planet emerges, Optimus Prime and the Autobots must team up with a powerful faction known as the Maximals. With the fate of humanity hanging in the balance, humans Noah and Elena will do whatever it takes to help the Transformers as they engage in the ultimate battle to save Earth.

Alex, Marty, and other zoo animals find a way to escape from Madagascar when the penguins reassemble a wrecked airplane. The precariously repaired craft stays airborne just long enough to make it to the African continent. There the New Yorkers encounter members of their own species for the first time. Africa proves to be a wild place, but Alex and company wonder if it is better than their Central Park home.